Bees, like all insects, have six sections to their legs: the coxa, trochanter, femur, tibia, metatarsus and tarsus. Each is connected by a joint and the one most like a knee is between the femur and tibia. ... Of all of these, the only one actually found in nature is the bee's knees, so perhaps that's what's so special.
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Do bees have knees and, if so, what's so special about them? - BBC Science